Princess Natty of Orchidhali is a member of the floral caste and lives in a luxurious, sunshine-filled palace covered. Had she not neglected her studies of sacred texts and tradition, she'd have learned that refusing all of candidates presented to her for marriage would result in her death! She is forced to flee her palace and ends up in the dark realm of the Untouchables, the very lowest caste of society, where a strange form of leprosy reigns. One of the Untouchables, Sami, doesn't seem to suffer as badly as the others. As Natty develops a friendship with Sami, she discovers that he owes his relative beauty to his activity as a mushroom trafficker. In exchange for the valuable fungi, Sami obtains time exposed to the sun: in fact, it is the complete absence of sun from the Untouchables' realm that leads to their deformed bodies and skin. Together, Sami and Natty make a plan to leave the shadows of the underworld and return to the light. But just as they are about to make their escape, a battalion of soldiers arrives...

This second and final volume of the Natty adventure picks up where it left off: Natty is alone in the underworld fighting for her survival while her family up above engages in a wild search for her. Sami manages to escape from prison, and thanks to their agility, strong will, and a dose of luck, he and Natty make their way back to the palace. After some initial difficulties, Natty is recognized by her parents as the Princess, and she and Sami live happily ever after. They even find a solution to bring sunlight to the lower levels of the kingdom.